Wetland vegetation in the sub(alpine) zone of the West Carpathians (Poland, Slovakia) was studied with particular reference to the following questions: (i) What are the main types of (sub)alpine fen, bog and spring vegetation above the timberline in the West Carpathians? (ii) Which major environmental gradients are associated with the variation in floristic composition? (iii) What determines the α-diversity of bryophytes and vascular plants in the different vegetation types? Vegetation plots were sampled and direct measurements of certain environmental characteristics recorded. Cluster analysis was used to distinguish the vegetation types, DCA and CCA to reveal the main vegetation gradients and environmental factors, and general regression models to identify the factors determining the α-diversity. Classification at the level of 12 clusters was ecologically and syntaxonomically interpretable. Two associations not mentioned in the most recent vegetation survey of Slovakia were distinguished and the syntaxonomical positions of others revised. The synthesis of collected and published vegetation data for the two countries has modified the classification concept of (sub)alpine wetlands in the West Carpathians. Whereas pH predominantly determined floristic differences among classes, different factors governed the species composition within classes. While the diversity of the vegetation of springs (Montio-Cardaminetea) was mostly determined by water chemistry, altitude and geomorphology appeared to be more important within fens and bogs. The species richness of spring vegetation was more influenced by mineral richness than water pH and for bryophyte richness the slope inclination was also important. The species richness of fens and bogs increased with pH. It is concluded that the ecological gradients influencing the floristic composition and species richness of (sub)alpine wetlands are strongly habitat-dependent.
Vegetation with species of Utricularia and that dominated by Eleocharis quinqueflora, which occupy the same habitats. was studied in minerotrophic mires and oligotrophic wetlands associated with ponds. Relative towater and soil chemistry, the communities of Utricularia ochroleuca s.l. and U. intermedia occurred in mineral-poor and those of U. minor and U. australis in mineral-rich conditions. Rare stands with U. vulgaris occurred in conditions that were intermediate in mineral richness. Four communities belonging to the class Isoëto-Littorelletea were distinguished. Vegetation without bladderworts and dominated by E. quinqueflora occurs in calcareous fens and belongs to the class Scheuchzerio-Caricetea fuscae (the Caricion davallianae alliance). Vegetation with U. intermedia is characterized by high vascular plant cover and belongs to the class Scheuchzerio-Caricetea fuscae. Utricularia ochroleuca s.l. prefers open, acidic and waterlogged depressions in peat, whereas U. intermedia grows mostly in the shade under vegetation canopy. In this study, U. minor and U. australis have been found mainly in the more alkaline and mineral-rich habitats, and both species also tolerated extremely high mineral richness.