Rubus perpedatus Žíla et H. E. Weber (sect. Rubus ser. Radula Focke) is described as a new species. It differs from R. radula Weihe in having distinctly pedate leaves, terminal leaflets with an emarginate base, longer pedicels with longer stalked glands and with many more prickles. It is distributed in the Bavarian Forest, the Bohemian Forest and in Upper Austria. An illustration, a list of herbarium specimens and a map showing the distribution of this new species are provided.
A new species of bramble, Rubus kletensis, of the section Corylifolii Lindley, series Sepincola (Focke) E. H. L. Krause occurring in South Bohemia and Upper Austria is described. The distance between the most distant localities exceeds almost 150 km. This distinct and relatively easily recognizable species grows in rather moist, eutrophic, synanthropic and sunny biotopes, and occurs most frequently in the vegetation of the class Galio-Urticetea, less frequently in that of the alliances Trifolion medii, Pruno-Rubion radulae, Sambuco-Salicion capreae, Berberidion and exceptionally in forest plantations and growths of pioneer saplings. A distribution map for this species and a list of all known localities are included, as well as a drawing of the species.
Rubus silvae-bohemicae is described as a new regional apomictic species belonging to the subgen. Rubus sect. Rubus ser. Micantes Sudre. It is a nemophilous bramble occurring in south-western and southern Bohemia (Czech Republic) and adjacent parts of Bavaria (Germany). It differs from the somewhat similar species, R. indusiatus Focke, by having no stellate hairs on the undersides of the leaves and a lower number of shorter stalked glands and lower number of prickles on first-year stems. An illustration of the new species (including a photograph of the type specimen), a list of localities and a distribution map are presented.
A new bramble species, Rubus silvae-norticae, section Rubus, subsection Hiemales E. H. L. Krause in Prahl, series Micantes Sudre, which occurs in S Bohemia, Upper Austria and Lower Bavaria, is described. It is recorded at 130 localities. The distance between the most remote localities is ca 100 km. The species grows most frequently in forest habitats (as a distinctly nemophilous ecoelement) such as ditches and edges of forest roads, plantations, forest margins and clearings. It mainly grows in mesic, acid and mineral-poor soils. Like, for example, R. clusii or R. ser. Glandulosi and unlike other relatively thermophilous Rubus species, it is able to grow and propagate itself at rather high altitudes, up to the mountain vegetation belt. The diagnostic characters that separate R. silvae-norticae from its most similar and sympatrically occurring species, R. clusii and R. muhelicus, are provided. In Austria R. silvae-norticae and some other brambles were mistakenly considered as R. helveticus, a bramble (probably a single biotype) described from Switzerland in 1870. The lectotype of Rubus helveticus is designated here and a photograph of the specimen presented. Also included is a distribution map of R. silvae-norticae, a list of revised herbarium specimens, a photograph of the type specimen and a pen drawing of the species. The significance of regional brambles for plant migrations and phytogeography is shown, based on the distribution of selected regional Rubus species occurring in the Czech and Austrian border area, which is a known mountain barrier to migration. The distribution patterns of the brambles support a theory about the routes of plant migration and the florogenetic connection between Austria and the Czech Republic. Rubus silvae-norticae, R. muhelicus and R. vestitus f. albiflorus are regarded as Danubian migrants (distributed from Upper Austria to S Bohemia), whereas R. gothicus s. l. (“south Moravian type”) and R. austromoravicus are considered to be Dyje-Kamp migrants (distributed from Moravia and Lower Austria to S Bohemia) within the Bohemian flora. Rubus kletensis is supposed to be a Vltava migrant within the Austrian flora (distributed from S Bohemia to Upper Austria).
The current state of the taxonomy of Rubus ser. Discolores in the Czech Republic is summarized. Since 1995, when the group was treated in the Flora of the Czech Republic (Holub 1995), six new species have been recognized, some of which are also known from adjacent areas. They are described in the present study: R. austroslovacus Trávníček, R. flos-amygdalae Trávníček et Holub, R. guttiferus Trávníček et Holub, R. parthenocissus Trávníček et Holub, R. pericrispatus Holub et Trávníček and R. portae-moravicae Holub et Trávníček. The first five belong to the group of triploid species close to R. montanus Lej. and R. grabowskii Weihe, whilst R. portae-moravicae is related to R. praecox Bertol. Two additional species, originally recognized under provisional names, were found to be identical to species described earlier: R. perperus H. E. Weber and R. phyllostachys P. J. Mueller; the latter is also found in Slovakia. At present, 17 indigenous species, one naturalized alien (R. armeniacus Focke) and one rare garden escape (R. ulmifolius Schott) of the ser. Discolores are known to occur in the Czech Republic. Distribution data and a key for the identification of all the species are presented.