Nutrient-rich terrestric habitats form small areas in the peaty alluvial plain of upper stream of the Vltava river. Their vegetation consists of birch and grey alder alluvial woodland, willow and bridewort scrub and tall grassland of sedges, grasses and forbs. A hypothesis that this vegetation is an Early Holocene relic is presented. The relict origin is supported by recent dynamics of habitats and vegetation, findings of palynology, palaeoecology and history of land use, and by the analogical composition and history of relict vegetation of northernmost Europe. The refugial effect of the habitat is suggested by stable conditions on high temporal and spatial scales, and by permanent reclaiming of open gaps along the stream.